Entirely developed and manufactured in-house, calibre 7135 operates at an exceptional 5 hertz, or 36,000 beats per hour, an unprecedented frequency for Rolex. This enables it to measure time to one tenth of a second. It also presents a particularly attractive design. Through the watch's transparent case back, the sparkle of the cut-out oscillating weight in yellow gold may be admired, as well as the delicate Rolex Côtes de Genève decoration on the bridges.

The most significant innovation, calibre 7135's Dynapulse escapement is a revolutionary alternative to the traditional Swiss lever escapement. Made primarily of silicon components and boasting an innovative architecture, this sequential distribution escapement is extremely lightweight, virtually insensitive to magnetic fields and significantly more energy-efficient than a traditional escapement.

Calibre 7135's oscillator features a balance staff whose material and manufacture presented a significant challenge, a balance wheel made from optimized brass, and a silicon Syloxi hairspring. The patented balance staff is made from a new high-technology ceramic exclusive to Rolex and is shaped by laser – a first for the brand. Optimized Paraflex shock absorbers give the movement heightened shock resistance in any position. Finally, thanks to the various materials used to produce the oscillator's components, it is able to withstand strong magnetic fields.

The Land-Dweller reinterprets the design of watches with integrated bracelets, characterized by a smooth continuous line between case and bracelet. Inspired by the emblematic Jubilee bracelet introduced on the Datejust in 1945, the Flat Jubilee bracelet has been specially designed for the Land-Dweller. Its links have a distinctive flat surface, a unique feature among our metal bracelets. It also comes with a whole new set of finishes, including polished centre links, and outer links with a technical satin finish on the top and polished sides. Viewed from the side, the polished chamfers on the top edges of the case and outer links form a circle of light, further accentuating the timepiece's harmonious design.

Surrounded by an angled flange, this graphic dial is adorned with a geometric decoration: a honeycomb motif developed using a high-technology laser. The satin finish on the intense white dial and the sunray finish on the ice blue dial are achieved using traditional techniques. On versions with a Chromalight display, the open numerals and the index hour markers, with luminescent material that extends the full length of the component, are exclusive to the range. They give the Land-Dweller optimum legibility in any circumstance. The hands, also luminescent, have been specially designed for the model. The seconds hand, in particular, stands out with the design of its counterweight, which adopts the dial's honeycomb motif.

Like all Rolex watches, the Oyster Perpetual Land-Dweller 36 and the Oyster Perpetual Land-Dweller 40 are covered by the Superlative Chronometer certification redefined by Rolex in 2015. This designation testifies that every watch leaving the brand’s workshops has successfully undergone a series of tests conducted by Rolex in its own laboratories according to its own criteria, following the official certification of the movements by the Swiss Official Chronometer Testing Institute (COSC). These in-house certification tests apply to the fully assembled watch, after casing the movement, guaranteeing superlative performance on the wrist in terms of precision, power reserve, waterproofness and self-winding. The precision of a Rolex Superlative Chronometer is of the order of −2/+2 seconds per day – the rate deviation tolerated by the brand for a finished watch is significantly smaller than that accepted by COSC for official certification of the movement alone.

The Superlative Chronometer status is symbolized by the green seal that comes with every Rolex watch and is coupled with an international five-year guarantee.